Understory

/ˈʌn.dəˌstɔː.ri/ noun

The understory is the layer of plants and small trees that grow under the main canopy of a forest. It is usually darker and more humid, with its own special plants and animals.

From "under" plus "story," using "story" in the sense of a level of a building. It imagines the forest as a multi‑level structure, with the understory as a lower floor.

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