Having calycules; a past-participle form indicating that an organism or structure possesses or is equipped with small cup-like appendages similar to a calycule.
The past participle form of the adjective/verb calyculate, using the -ed suffix. This alternate form (alongside calyculate) appears in older botanical texts and scientific descriptions.
The fact that botanists used both 'calyculate' and 'calyculated' to mean the same thing shows how scientific language was still settling its grammar rules in the 1800s—there wasn't one official police force for terminology!
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