In a manner that involves chuckling or quiet laughing; with gentle, suppressed amusement.
From chuckle with the adverbial suffixes -ing and -ly, creating an adverbial form that emphasizes the manner of action.
This adverb appears frequently in 19th-century literature to describe how characters respond to mild humor—it's far more specific than 'laughingly' and carries a sense of warmth rather than loud amusement.
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