A rooster or cock; a bird that crows at dawn.
From cockcrow (the sound and time) + -er (agent suffix). This term evolved in Middle English as a way to describe the animal responsible for making the characteristic dawn sound.
While we might call this bird a 'rooster' today, 'cockcrower' is a delightfully descriptive name that emphasizes its actual function—it's literally 'the thing that crows at cockcrow.' It's the kind of practical, onomatopoetic naming that makes Old English feel very logical.
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